Mushroom-Term Glossary
Acanthophysis -- see hyphidium
Aciculate -- Very slender, with sharp top, therefore needle-shaped
Acidulous -- Slightly acid.
Acrid -- Taste burning or peppery, in this program the description of 'acrid' is rendered as 'peppery'
Acula (plural aculae) -- Spine.
Aculeate -- Of cystidia, tapered so that only the very basal portion is relatively swollen, the entire cystidium being shaped like a spine, therefore spine-shaped; of spore, means having narrow spines
Acuminate -- Gradually narrowed to a point
Acute -- Pointed, sharp; less than a right angle
Adnate -- Pertaining to the attachment of the fertile tissue (the gills, tubes, spines, etc.) to the stipe of the fungus in which the attachment is typically perpendicular into the stipe, i.e. without dipping towards the pileus or down the stipe.Adnexed -- Pertaining to the attachment of the fertile tissue (the gills, tubes, spines, etc.) to the stipe of the fungus in which the fertile tissue typically curves upwards towards the pileus of the fungus before attaching to the stipe.
Agaric -- A term commonly used to describe a fungus having a cap (pileus), gills (lamellae), and a stem (stipe), i.e., what most people would call a mushroom.
Agaricologist -- A person who studies gilled mushrooms
Agglutinated -- Surface fibrils or scales drawn together in clumps
Allantoid -- Sausage-shaped, tubular and slightly curved with rounded ends
Alliaceous -- Smelling or tasting like onions or garlic
Alveolate -- Surface of cap or spore with broad pits
Amanitin -- Same as amatoxin.
Amanitoid -- Like Amanita, with free or slightly adnexed gills, a volva, and a ring
Amatoxin -- Cyclic peptide found in Amanita and other genera that are very toxic
Ampullaceous -- Flask-shaped
Amygdaliform -- Almond-shaped
Amyloid -- A chemical staining reaction in which the tissue, spore wall ornamentation, etc. stains bluish-black in Melzer's reagent. Examples include the spore ornamentation of species in the genera Russula and Lactarius.Anamorph -- The asexual reproductive manifestation of a fungus, characterized by asexual spores
Annular -- Resembling a ring or referring to a ring, as in an annular zone on stem
Annular zone -- a band of fibrils or gluten around stem, often becoming darkened by spores, normally derived from veil remnants, but too obscure to be a ring
Annulus -- A ring of tissue on the stipe of mushroom formed by the rupture of a membrane (the partial veil) connecting the cap and stipe of a developing mushroom. A special layer of tissue that connects the margin of a mushroom pileus to the stipe that can either form a ring around the stipe, hang as fragments from the margin of the pileus, or be variations of the two. Examples may be found in many genera such as Amanita, Cystoderma, Lepiota, and Suillus, among others.


Anthesis -- Point of development of fruiting body at which the fresh unexpanded cap is in "full flower", contains the features for identification, and is at the brink of spore release
Apex -- Top, highest part
Apical -- Near top
Apical pore -- Same as germ pore, not to be confused with apiculus, which is the other end of the spore
Apiculus -- Nipple-like projection; nipple-like projection on spore which corresponds to the area that was attached to the basidium, sometimes used to refer to a projection on the other end of the spore, same as hilar appendage and not to be confused with apical pore (germ pore)
Appendiculate -- Margin of cap fringed with hanging fragments of the veil; (of cystidium) having an appendage; (of a spore) having one or more setulae
Applanate -- Horizontally expanded, plane, flat
Arachnoid -- Cobweb-like
Arcuate -- Forming an arch; of gills, means that the middle of the lower edge of the gill is higher than its ends
Areolate -- Cracked in age, The cap surface of just about any mushroom can become cracked in dry weather conditions, but some species typically develop cracked caps in normal weather conditions.

Armillarioid -- With attached gills, fleshy stem and ring
Ascending -- Refers to gills that curve upwards from the margin of the cap to the attachment at the stem, as in conic or unexpanded cap
Ascus -- A specialized sexual reproductive cell found in the fertile area of the hymenium of all Ascomycetes. An ascus is typically club shaped and which forms internally 4 or 8 ascospores, usually in a row.Ascomycetes -- The Ascomycetes are fungi that produce their spores in little tubes called "asci" (the singular is "ascus"). The tubes are located on the spore-bearing surface of the mushroom. There are no mushrooms with gills, pores, or spore-bearing spines among the ascos. Morels, False Morels, and Helvella, and species of cup Fungi are "ascos."
Ascomycota -- Pylum that includes the largest group of fungi, those that produces their spores in sacs called asci, but does not include any gilled mushrooms
Asperulate -- Of spores, appearing roughened with tiny points; or roughened with small warts
Asterostromelloid -- Type of pellis found in Resupinatus composed of swollen terminal elements with short approximately perpendicular branches
Astringent -- Causing a contraction or pucker of the mouth membranes
Atomate -- A powdered surface consisting of minute shiny particles
Attenuate -- Gradually narrowed
Avellaneous -- Dull grayish brown, hazel-brown, or light gray-yellow-brown, or closer to drab, or gray tinged with pink
Azonate -- Without zones, without concentric markings
Azure -- Sky Blue in color
Badious -- Dark red brown
Baeocystin -- An indole alkaloid (4-phosphoryloxy-N-methyltryptamine), closely related to and often found with psilocybin, possibly hallucinogenic with comparable effect to psilocybin
Bald -- No warts or hairs or raised scales, fibers or patches, same as glabrous and as used here equivalent to naked
Basal -- Near the base
Basal mycelium -- Fungal cells at the base of the stem, ranging from a few fibrils to a velvet layer
Basal tomentum -- Same as basal mycelium
Basidiocarp -- Fruiting body: the whole reproductive structure of a mushroom including cap, gills, and stem
Basidiole -- Immature basidium
Basidiome -- Fruiting body: the whole reproductive structure of a mushroom including cap, gills and stem
Basidiomycete -- Fungus belonging to Basidiomycetes
Basidiomycetes -- Class that includes most gilled mushrooms as well as chanterelles, tooth fungi, boletes, polypores, puffballs, bird's nests, jelly fungi etc.
Basidomycota -- Phylum that includes classes Basidiomycetes (includes most gilled mushrooms as well as chanterelles, tooth fungi, boletes, polypores, puffballs, bird's nests, jelly fungi etc.), Teliomycetes (rust fungi etc.), and Ustomycetes (smut fungi etc.)
Basidiospore -- Sexual spore, the usual spore produced on gills
Basidia -- The plural form of basidium. Basidia cannot be seen with the naked eye. They cover the gills of gilled mushrooms, and the interior surfaces of the tubes in mushrooms with pores.Basidium -- A specialized sexual reproductive cell found in the fertile area of the hymenium of all Basidiomycetes, typically shaped like a baseball bat. A basidium possesses four slightly inwardly curved horns (sterigma) to which the basidiospores are attached.
Basionym -- The earliest name of a taxon on which a later name has been founded
Bay -- Red-brown approaching but lighter than chestnut
Beige -- Light grayish yellowish brown
Bell-Shaped -- In the shape of a bell (like the
Benzo Brown -- Pale purplish brown, dark brownish, violaceous fuscous, in Ridgway 1912 gray brown with a pinkish tinge
Bibulous -- Of surface of cap, capable of absorbing moisture
Bicolorous -- Of Two colors
Bifurcate -- Divided into two branches of equal or unequal length
Bilateral -- Of gill tissue, same as divergent
Binding Hypha -- Thick-walled, narrow, highly branched non-septate hypha found only in trimitic tissue
Binomial -- Blackish brown, a warm dark brown color, like sepia, dark yellow-brown
Bister -- Blackish brown, a warm dark brown color, like sepia, dark yellow-brown
Bloom -- Minutely velvety or powdery surface
Bolete -- The common name for soft textured fungi which generally have pores instead of gills
Boletinoid -- Pore Surface
Bone Brown -- Dark vinacious brown
Brachybasidiole -- Cell within the hymenium, which unlike the clubshaped basidioles enlarges to be round or elliptic in shape
Brittle -- Breaking easily, rigid and breaking with a snap; of stem, forms a sharp non-fibrous edge when broken
Broad -- When used of gills, refers to the height (depth) of the gill, which may be narrow, moderately broad or broad
Broadly Convex -- Of cap, convex but mostly flattened apart from downcurved margin, same as plano-convex
Broad-leaved -- Of trees that have leaves rather than needles
Brown Rot -- Carbonizing decay (cellulose-composing decay)
Brownish Drab -- Purple gray with slight brownish cast
Bruises -- Sometimes described the stem or the gills/pores may bruise on some species when handled.



Bryicolous -- Growing on or among mosses
Buckhorn Brown -- Dull yellowish brown
Buff -- A pale yellow toned with gray-brown, i.e. a dingy yellowish brown or very pale tan
Bulb -- A part shaped like the underground part of an onion or daffodil or similar plant
Bulbose -- Having a bulb or bulging area; of stem, with an enlarged base
Bulbous -- Refers to a bulbous like swelling at the base of the stem (stipe), often underground
Burnt Sienna -- Watery strong red-brown, dark orange brown
Butt Rot -- A rot confined to base or roots of a tree
Button -- Young fruiting body before it has opened up. An immature specimen. See Photo Below.

Byssoid -- Of mycelium, the condition when fine filaments spread from the base of the stem or fruiting body over substratum
Caesious -- Pale bluish-gray
Caespitose -- growing in close groups or close clusters or tufts (may be from a common base, but stems not joined together), see clustered
Calcareous -- with a hood; of spores, the outer layer separating to form a partial envelope or bag around spores, often with blisters or loose areas, as in some Galerina
Cambium -- the cellular plant tissue, or tissue layer, responsible for the increase in girth of stems and roots
Campanulate -- bell-shaped
Canaliculate -- of stem, furrowed or fluted
Candidous -- shining white
Canescent -- becoming densely covered with whitish or grayish down or becoming gray or hoary
Cap -- See pileus. The pileus which is the umbrella or bell like 'hat' of the mushroom. The pileus holds the spores in either gills or pores. Caplike part of fruiting body which supports the gillsCarpophore -- The complete fruit body of the fungus (i.e. cap, stem, gills, etc). Sporophore and sporocarp are other names also used
Capillitium -- modified threadlike branched hyphae in the spore mass of many Gasteromycetes
Capitate -- with a head or cap, abruptly enlarged at top
Capitellum -- small head
Capitulum -- a relatively small, spherical head or ball carbonizing decay a decay characterized by loss of cellulose and pentosans, leaving lignon in the form of a brown carbonous mass
Carinate -- of spores, furnished with a keel, boat shaped cartilaginous of stems: firm, tough and pliant (flexible), typically under 5mm in diameter at top of stem; having the consistency or appearance of cartilage; sometimes used even of fragile stems and implying brittle, not pliant
Cartridge buff -- pale yellowish
Catenate -- in chains or end to end series
Catenulate -- in chains or end to end series
Caulocystidium (plural caulocystidia) -- a sterile cell located on the stem
Cespitose -- Clustered mushrooms fused at the base. Also spelled caespitose. Growing in tufts or close clusters from a common base, but not grown together. See Photo Below.

Cellular -- composed of rounded cells, not threadlike ones
Cellulose -- a component of wood and plant cell walls made of glucose units
Ceraceous -- waxy
Chamois -- pale yellow
Chartreuse -- a color between green and lemon yellow, apple green, the color of Letharia (wolf lichen)
Chateura drab -- pale fuscous
Cheilocystidium (plural cheilocystidia) -- a sterile cell located on the edge of a gill
Cheilopseudocystidium (plural cheilopseudocystidia) --pseudocystidium located on the edge of a gill
Chestnut -- dark red-brown
Chlamydospores -- thick-walled asexual spores formed by breaking up of hyphae, that served to survive adverse conditions
Chrome-yellow -- intense or strong yellow; also can be deep orange or even reddish orange
Chrysocystidium (plural chrysocystidia) -- a type of cystidium that is highly refractive in once-dried tissue revived with a potassium hydroxide (KOH) solution, appearing as a yellowish brown shapeless mass within the cell
Cigar -- brown a dark blackish brown, like that of the tobacco found in a cigar
Ciliate -- having a fringe of hair-like ciliae; appearing fringed
Cinereous -- ash-colored, dirty white
Cinnabar -- deep bright rather orange red
Cinnamon -- a light brown with a little pink
Cinnamon-buff -- pale dingy tan
Cinnamon-drab -- violaceous brown with only a faint cinnamon tint, in Ridgway 1912 pinkish gray brown
Cirrate -- rolled round (curled) or becoming so
Citrine --a light greenish yellow
Citron yellow -- greenish yellow
Clamp -- same as clamp connection
Clamp connection -- small tubular elbow-like bypass across the walls between fungal cells, used to distinguish genera and species, functionally providing a bypass for one of two nuclei to insure their equal distribution in new cells. A special connection which forms at the junction of two adjacent fungal filamentous cells. A clamp connection looks something like the handle on a coffee cup. However, it may be flattened against the wall of the cells or may have a large opening (in this case a keyhole clamp) that allows the migration of nuclei between developing cells.
Class -- classification group above order but below phylum: suffix for the fungi is -mycetes
Clavate -- like a caveman's club; when used of stems, implies base is thicker and stem tapers upward; when used of cystidia, implies part that extends outward beyond the hymenium is thicker, same as club-shaped
Clay color -- clay color, resembling dull ochraceous-cinnamon-brown, in Ridgway 1912, closer to ocher or bright yellow brown
Clitocyboid -- resembling in general form a mushroom of the genus Clitocybe, typically with decurrent gills, fleshy-fibrous stem, and without a ring or volva
Close -- of gill spacing, nearly touching but with visible space between, intermediate between crowded and distant, the order being crowded, (subcrowded), (subclose), close, subdistant, distant
Club-shaped -- like a caveman's club; when used of stems, implies base is thicker and stem tapers upward; when used of cystidia, implies part that extends outward beyond the hymenium is thicker, same as clavate
Clustered -- growing together, either very close or from a common base
Cochleariform -- spoonlike in form
Collybioid -- resembling in general form a mushroom of the genus Collybia, typically with expanded caps (convex to broadly convex to flat) often with downcurved to incurved margin, cartilaginous or brittle stems not more than two or three times in length the diameter of the caps, without ring
Colors -- Always in Latin to be more specific and scientific, they are as follows:
aeruginose (verdigris-green, (malachite-green), the color of oxidized copper)
alutaceous (light leather colored, usually interpreted as light tan or medium yellow brown)
argillaceous (clay color, resembling ochraceous-cinnamon-brown)
avellaneous (pale pinkish gray);
azure (Sky Blue in color)
badious (Dark red brown)
bay (Red-brown approaching but lighter than chestnut)
beige (Light grayish yellowish brown)
benzo brown (Pale purplish brown, dark brownish, violaceous fuscous, in Ridgway 1912 gray brown with a pinkish tinge)
binomial (blackish brown, a warm dark brown color, like sepia, dark yellow-brown)
bister (blackish brown, a warm dark brown color, like sepia, dark yellow-brown)
bone brown (Dark vinacious brown)
buckhorn brown (Dull yellowish brown)
buff (A pale yellow toned with gray-brown, i.e. a dingy yellowish brown or very pale tan)
burnt sienna (Watery strong red-brown, dark orange brown)
fulvous (rusty brown or tawny);
fuscous (grayish or grayish brown);
isabella (yellowish brown to light olive brown);
isabelline (yellowish brown to light olive brown);
livid (grayish or bluish gray);
ochraceous (brownish yellowish);
olivaceous (olive--the greenish color)
rubescent (blushing reddish or pinkish);
Columella -- a sterile column of tissue projecting into the spore mass of Gasteromycetes
Complex -- a cluster of taxonomically related similar species typified by a particular species, as in Conocybe tenera complex
Compressed -- of a stem, elliptical to flattened in cross section
Concentric -- having rings or circular zones
Conchate -- like an oyster shell
Concolorous -- having the same color
Confluent -- going towards the same point
Conglobate -- massed into a ball; (of the bases of stems), together making a fleshy mass
Conic -- shaped like a cone
Conic-campanulate -- of cap, bell-shaped with conic umbo
Conic-convex -- of cap, convex with conic umbo
Conidium (plural conidia) -- asexual fungal spores formed by the pinching off of hyphae
Conifer -- one-bearing tree
Connate -- of stem, joined by growth, e.g. when stems of several sporocarps are joined together, see clustered, caespitose
Connivent -- (of margin of cap) converging on the stem
Conspecific -- belonging to the same species
Context -- flesh of cap or stem (excluding the surface layer)
Convergent -- of gill hyphae, projecting inward and downward away from cap as seen in cross-section
Convex -- regularly rounded, domed, like an inverted bowl
Convex-campanulate -- of cap, bell-shaped with flattened or convex center
Convex-depressed -- of cap, convex with depressed center
Coprophilous -- growing on dung
Coriaceous -- leathery in texture
Cortina -- a web-like or silky veil extending from the cap margin to the stem in young mushrooms of certain species, soon disappearing or leaving remnants on stem or cap margin. A special type of annulus that is filamentous, resembling a spider web, attached from the margin of the cap (pileus) to the stem (stipe) when young. In age only a few fibers may remain on the cap margin or the stipe.


Cortinarioid -- having general shape of Cortanarius; applied to any mushroom with attached gills and cortinate or fibrollose partial veil
Cortinate -- with a cortina, weblike
Crawshay -- mycologist who designated color types for Russula spores, ranging from white (A) to deep ochre-orange (H)
Cracked -- surface having split in some way
Cream-buff -- a moderately dark buff
Crenate -- with notched edge or rounded teeth, scalloped
Crenulate -- finely scalloped
Crimson -- rich deep red inclining to purple
Crisped -- of gills, finely wavy
Crowded -- of gill spacing, very close, touching or with almost no space between, the order being crowded, (subcrowded), (subclose), close, subdistant, distant
Cucullate -- like a top hat; like a cowl or hood
Cuneate -- wedge-shaped
Cupulate -- cup-shaped
Cup -- The description given to the saucer shape of the Ascomycetes group
Cystidium -- A specialized sterile end cell formed anywhere in fungal tissue. It is most commonly found in the hymenial layer of tissue, but may also be found on the surface of the cap, the surface of the stipe, or even within the sterile tissue of the stipe. There are many different types of cystidia; they are named based on the location where they are found, e.g. Dermatocystidia- on the surface tissues; Pileocystidia- found on the surface of the pilius; Caulocystidia- found on the surface of the stipe; Cheilocystidia- on the edge of the gill; Pleurocystidia- on the face of the gill; Endocystidia- form in the tramal tissue of the cap, or stipe; OR on their morphology, function, chemical reactions etc. such as Leptocystidia-which are thin-walled, smooth and do not have distinctive contents and are not tramal in origin; Gloeocystidia- which are variable in shape and stain easily or have conspicuous contents; Lamprocystidia- which are thick-walled and without conspicuous contents, etc.Cuspidate -- sharp, pointed
Cuticle -- the cap skin or surface layer of cells; same as pellis, and thought by some to be incorrectly used in this situation as it refers in botany to the waxy surface of certain leaves
Cutis -- type of "cuticle" (pellis), surface layer of cap cells if they are differentiated from the underlying tissue and arranged more or less parallel to cap surface: if the hyphae are radially arranged, it is a parallelocutis; if they are not, it is called a mixtocutis; the prefix ixo- can be added to indicate that elements are gelatinized; see also pileipellis, derm
Cyanophilic -- said of basidia, spores or hyphae whose walls absorb a greater amount of cotton blue than their contents; said of contents of cystidia that absorb cotton blue
Cylindric -- of the same diameter throughout its length; of stem, terete (not compressed); of spores, according to one set of criteria ratio of length to width 2-3: less would be oblong, more would be bacciliform
Cyphellaceous -- refers to groups of basidiomycetes, often small and disk- or cup-shaped, which have been associated with the family Cyphellaceae: many of them have been redistributed to other families
Cystidium (plural cystidia) -- a sterile cell frequently of distinctive shape, at any surface of a fruiting body, (sometimes used in a strict sense restricted to sterile cells in hymenium), classified by 1) position: pileocystidium (cap), pleurocystidium (gill face), cheilocystidium (gill edge), caulocystidium (stem) etc., 2) form: leptocystidium (smooth, thin-walled, without discernible contents), lamprocystidium (thick-walled), metuloid (thick-walled encrusted; or a type of lamprocystidium which is rounded at top or with a variable shaped top), etc., 3) contents: chrysocystidium (like leptocystidium but with highly staining contents), gloeocystidium (thin-walled, usually irregular, contents colorless or yellowish and highly refractile) etc., 4) origin: pseudocystidia (derived from a conducting element, oily contents), macrocystidium (arising deep in the flesh of Lactarius or Russula); 5) often further described by shape: aciculate (needle-shaped), aculeate (spine-shaped), ampulliform (flask-shaped), capitate (with a head), clavate (club-shaped), cylindric, fusoid (spindle-shaped), furcate (forking), lageniform (gourd-shaped), lanceolate (lance-shaped), lecythiform (bowling pin shape), napiform (turnip-shaped), pyriform (pear-shaped), rostrate (with a beak), subulate (awl-shaped), tibiiform (like a tibia), turbinate (top-shaped), urticoid (with a swollen base and a long gradually narrowed apex), utriform (bladder-shaped), ventricose (wider in middle), vesiculate (bladder-shaped)
Cystoderm -- applied to pellis consisting of rounded elements; same as polycystoderm
Daedaloid -- with elongated and sinuous (curving) openings
Deciduous -- referring to trees that seasonally shed their leaves; or referring to anything that falls off, such as granules that tend to fall off the cap
Decorticated -- of dead wood without the bark
Decumbent -- (of stem) with the lower end lying against the substratum
Decurrent -- refers to gills that run down the stem: i.e. attachment at stem is wider than average height of gill. Pertaining to the attachment of the gills to the stipe, in which the gills curve partly down the stipe towards the base of the stipe.
Decurved -- referring to a cap margin or scales means curved downward
Deflexed -- same as decurved
Delignifying decay -- a lignin and cellulose decomposing rot, leaving the wood light colored and fibrous
Deliquesce -- melt into liquid, usually referring to the gills and cap of Coprinus or of some species of Bolbitiaceae
Deliquescing -- The process by which gills in the genus Coprinus rapidly break down into a black ink-like liquid, droplets of which disperse spores.
Deliquescent -- melting into a liquid
Dendrophysis -- see hyphidium
Denticulate -- finely toothed, or lined with triangular fragments of tissue
Depauperate -- undeveloped because of lack of favorable conditions
Depressed -- of cap, having the middle lower than the edge; of gills, sinuate; depressed adnate refers to an adnate gill with a portion of the gill lower than its outer edge
Derm -- surface layer of cap cells if they are differentiated from the underlying tissue and arranged more or less perpendicular to cap surface: if the elements are a single row or roundish cells, this is a cellular derm; if cells are elongated and all reach the same level, this is a palisoderm; if cells are elongated and of different lengths, it is a trichoderm; the prefix ixo- can be added to indicate that elements are gelatinized
Dermatocystidium (plural dermatocystidia) -- sterile cell (cystidium) on the cap surface
Dextrinoid -- staining yellowish brown or reddish brown in Melzer's reagent. A chemical staining reaction in which the tissue, spore wall ornamentation, etc. stains reddish to reddish-brown in Melzer's reagent.
Diaphanous -- transparent or nearly so
Dichophysis -- repeatedly dichotomously branched modified terminal hypha in spore-bearing surface
Dichotomous -- repeatedly dividing or forking in pairs
Differentiated -- developed so as to be different from surrounding cells; of cystidia, distinguishable from surrounding cells
Dimidiate -- of cap, semicircular; of gills, that reach only halfway to stem
Dimitic -- consisting of 2 kinds of hyphae, such as generative and skeletal hyphae, or generative and binding hyphae, see also sarcodimitic
Dingy -- color appearing grimy or dirty
Disc -- center of the cap
Discoid -- dish-shaped
Distant -- of gill spacing, meaning the gills are spaced far apart, the order being crowded, (subcrowded), (subclose), close, subdistant, distant
Divergent -- of gill hyphae, projecting outward and downward away from cap as seen in cross-section. Usually referring to gill trama, in which the tramal hyphae branch outward from the gill center towards the hymenium and downward towards the gill edge.
Diverticulate -- said of hyphae or cystidia that have numerous peg-like protuberances or small branches scattered over their surfaces
Division -- classification grouping equivalent to phylum for plants and fungi
Double -- of ring, composed of two distinct layers of tissue, the lower layer typically being cottony or fibrillose
Downy -- with fine soft hairs
Drab -- a dull medium or brownish gray, dark gray with shades of yellow; gray with violet overtones; in Ridgway 1912, a gray-brown
Drab gray -- brownish gray
Dry -- surface not sticky or slimy or hygrophanous, feeling as if there is no moisture on surface
Dull -- opposite of shiny
Duplex -- flesh of two distinct textures
Dusky -- somewhat dark, implies absence of light and color
Eburneous -- white, like ivory
Eccentric -- off center; of stem attachment, attached away from center of cap but not at its edge
Echinate -- having sharply pointed spines, like a sea-urchin
Echinulate -- Referring to spiny ornamentation; e.g. the fine spines seen on the surface of some species of puffballs in the genus Lycoperdon. Also, the finely spined spores of Laccaria species. See Photo Below.

Ecru -- light brown or the color of unbleached linen
Ecru drab -- brownish gray with slight pinkish cast
Ectomycorrhiza -- system in which mycelium branches through soil and forms a covering around individual rootlets, growing between the outer rootlet cells, exchanging phosphorus for compounds that the plant produces by photosynthesis, formed in trees of the family Pinaceae and Fagaceae; Genera which are predominantly ectomycorrhizal include Amanita, Cortinarius, Gomphidius, Hebeloma, Hygrophorus, Inocybe, Laccaria, Lactarius, Leucocortinarius, Naucoria, Paxillus, Rozites, Russula, Tricholoma
Ectotrophic mycorrhiza -- same as estomycorrhiza
Effused -- spread out over material it is growing on without regular form
Effuso-reflexed -- resupinate patch whose edges are bent back to form a cap
Egg -- Young stage of a stinkhorn or other mushroom before it actually buds open. See Photo below of an Amanita egg.

Elastic -- springing back to its original shape
Elevated -- of cap margin, describes cap with margin uplifted, usually seen in age
Ellipsoid -- like an oblong sphere, often referring to the three dimensional shape of a spore
Elliptic -- like an oblong circle, often referring to the outline (as opposed to the three dimensional shape) of a spore, according to one set of criteria, ratio of length to width is 1.15-1.60
Elongate -- of spores, same as oblong, at least according to one definition, but may refer to cylindric as well
emarginate -- of gills, with a notch near stem, Largent & Baroni equate it with abruptly adnexed, but Ainsworth's Dictionary of the Fungi appears to equate it to sinuate (notched at the proximal end at junction with stem), and Hansen illustrates it as a deeper notch of the sinuate type; also used to describe a particular kind of bulb on stem
Encrusted -- same as incrusted
Endomycorrhiza -- system in which mycelium branches through soil and grows between and within root cells, exchanging phosphorus and other nutrients for compounds that the plant produces by photosynthesis, formed in 90% of seed plants and conifers, except conifers of the family Pinaceae; Some Armillaria species form endomycorrhiza with orchids
Endosporium -- the innermost layer of the spore wall
Endotrophic mycorrhiza -- same as endomycorrhiza
Entire -- with even edges: of gills, not serrated or toothed; of cap margin, means the outline is smooth and uninterrupted (not lobed, or irregular, crenate or serrate
Epicutis -- the outermost layer of the pellis ("cuticle"), properly called the suprapellis
Epigeous -- growing on or above ground
Epiphytal -- growing on leaves
Episporium -- the layer of the spore that is usually the thickest, giving the spores their form and rigidity, outside the endosporium and inside the exosporium
Epithelium -- a type of cellular pellis in which elements are in chains so that the pellis is many-layered, same as polycystoderm
Equal -- of a stem, the same diameter throughout its length, cylindric; of gill, broad (high) to same extent throughout length or alike in length
Ericaceous -- of the heath family of plants
Eroded -- of the margins of cap or gills, developing irregular jagged edges as a result of deterioration, irregularly
Erumpent -- fruiting body erupting through ground but barely rising above it
Esculent -- edible
Evanescent -- soon disappearing, fleeting. Rapidly disappearing.
Even -- of cap margin, means not wavy or lobed: the bottom line of the margin as seen from the side is a single flat plane revolving around the stem; of gill edges, means not toothed, eroded, fringed etc; of surface of cap, stem or spores means without striations, elevations or depressions
Ex -- from, first published validly by second author
Excrescence -- of cystidia, outgrowth surface
Exosporium -- the layer of the spore that becomes the outermost when the perisporium disappears, outside the episporium
Expanded -- cap fully developed; cap spread out
Expanding -- of cap, spreading out
Fabaceous -- bean-like
Face -- of gills, the side as opposed to the edge (margin). The side of a gill
Face view -- of spores, when the basidium with attached spores is viewed from the side, the spores directly above and in the center are being viewed in face view and the ones at the sides are in profile view, when not attached to basidium the shape that is wider will generally be the face view if there is a difference
Facultative -- referring to an optional situation, sometimes, not necessarily; not obligate; of a parasite, able to live also as a saprobe (on dead material or from substances produced by living ones)
Fairy Ring -- Click Photo for detailed story and description. A circle or arc of mushrooms
Falcate -- of spores, sickle-shaped
Falsely echinulate -- of spores, appearing ornamented with fine spines in optical section but actually with smooth outline, as in Fayodia
Family -- a classification group above genus and species, but below class and order, suffix is -aceae
Farinaceous -- of odor, with the smell of fresh ground meal from whole grain, especially wheat, same as mealy. An odor variously described as that of raw potatoes, raw cucumbers, or even of soaps; mealy.
Fasciated -- of stems, caps, grown together so that tissues are intimately continuous, compare caespitose
Fascicle -- a little group or bundle of fruiting bodies, hyphae or cystidia
Fasciculate -- of fibrils, scales, stems, crowded in bundles
Felted -- same as felty
Felty -- appearing like felt, covered with more or less erect short fibrils which are long enough that some collapse; densely matted and interwoven, same as matted-fibrillose
Ferruginous -- rusty red
Fetid -- ill-smelling
Fibril -- thin thread-like fiber
Fibrillose -- composed of delicate fibers which are long and evenly arranged on the surface. Possessing surface fibrils.
Fibrous -- composed of tough, stringlike tissue
Filamentous -- composed of hyphae (threadlike cells). Composed of thread-like cells.
Filiform -- threadlike, long and slender
Fimbriate -- fringed
Finely adnexed -- of gills, so narrowly attached that they give the appearance of being free
Fistulose -- of stem, having the flesh empty of fibrils, same as hollow, tubular
Flabelliform -- fan-shaped
Flaccid -- not firm, flabby
Flaring -- spreading out
Flat -- of cap, the margin being on the same level as the center, same as plane and applanate
Fleeting -- quickly disappearing, used here as equivalent to evanescent or fugacious
Flesh -- the tissue of cap or stem, not including the surface
Fleshy -- soft as opposed to tough; having significant substance
Flexuous -- of the stem, or of cystidia, bent alternately in opposite directions
Floccose -- with easily removed cottony or woolly tufts; woolly or cottony; dry and loosely arranged; having the appearance of cotton flannel. Having a cottony appearance. Seen in some species of Amanita.
Flocculose -- with fine, easily removed cottony or woolly tufts; finely woolly or cottony
Fluted -- of cap, with radial ridges; of stem, with longitudinal ridges
Foetid -- ill-smelling
Foray -- a field trip to hunt mushrooms, such as the term "foraging"
Foray list -- a list of species compiled from a field trip, often from macroscopic features, and often not as accurately as in geographic listings found in monographs or journal articles
Forked -- of gills, dividing into two or more branches as they goes away from stem
Forking -- of gills, dividing into two or more branches as they goes away from stem
Form -- a consistent appearing variation of a species, with less variation than a variety, often not sufficiently hereditary as to characterize homogeneous populations
Forma -- see form
Free -- refers to gills that are not attached to stem
Friable -- crumbling easily
Fringed -- with a border of parallel threads or fibers, so that the edge is somewhat jagged and not smooth
Frondose -- of a forest or the wood of hardwood trees
Fruit-body -- same as fruiting body
Fruiting body -- the whole reproductive structure of a mushroom including cap, gills and stem
Fugacious -- soon disappearing, fleeting
Fuliginous -- sooty brown or dark-smoke colored
Fulvous -- fox-colored, deep orange to reddish orange, reddish cinnamon brown
Fungus (plural fungi) -- an organism that lacks chlorophyll, consists of filamentuous tubular branching cells with nuclei, and reproduces by spores
Funnel-shaped -- with a very deep depression, like that of a funnel
Furcate -- forking; forked or branched irregularly
Furfuraceous -- scurfy, surface covered with branlike particles resembling scales, coarser than granular
Fuscous -- color of a very dark storm cloud: variously described as combinations of gray, brown, purple, or black
Fuscous-black -- a dark, dusky black or a black with a dark reddish-gray component
Fuscous-brown -- a dark, dusky brown or a brown with a dark reddish-gray component
Fuscous-purple -- purple with a dark reddish-gray component or somewhat dark (dusky) purple
Fusiform -- spindle-shaped, narrowing from middle to both ends
Fusoid -- somewhat spindle-shaped
Fusoid-ventricose -- of cystidium, tapered toward both ends but distinctly enlarged in the middle
Gasteromycete -- a basidiomycete that does not actively discharge it spores, formally constituting a class of basidiomycetes
Gastroid -- not forcibly discharging its spores
Gelatinous -- jelly-like in consistency or appearance; applied to tissue whose hyphae become partially dissolved and glutinous in wet weather and when mounted in water under the microscope appear more transparent and wider, loosening from one another
Generative hypha -- thin-walled branched hypha
Geniculate -- bent (like a knee)
Genus (plural genera) -- classification grouping below family but above genus: first letter is given in upper case
Genus -- Taxonomic term meaning a group of similar species. Genera which are closely related are placed into families
Germ pore -- a soft spot in the wall of certain spores, through which the fungus first starts to grow, same as apical pore (but not the same as apiculus)
Gibbous -- of cap, with an unsymmetrical convexity or umbo, or with convexity on one side
Gill -- gill, the spore-bearing platelike structure extending underneath and from the center of the cap like a spoke of a wheel
Gill Attachment --
Adnate -- Pertaining to the attachment of the fertile tissue (the gills, tubes, spines, etc.) to the stipe of the fungus in which the attachment is typically perpendicular into the stipe, i.e. without dipping towards the pileus or down the stipe. See Below Photo.


Decurrent -- Gills running down the stem. See Below Photo.



Free -- Gills not directly attached to the stem(stipe). See Photo Below. Common in Amanitas.



Gill Spacing --
Distant-- Subdistant-- Close-- Crowded--








Gilvous -- a bright yellowish brown, yellowish leather colored
Glabrate -- becoming glabrous, becoming bald
Glabrous -- bald, without hairs or raised fibers or scales or raised patches
Glandular -- with sticky drops or glands
Grandular Dots -- Glandular dots appear on the stems of some mushrooms, commonly in the genus Suillus. The dots are usually very small, and result from clusters of pigmented, inflated cells on the stem surface. See Photo Below.

Glaucous -- sea-gray; sea-blue-green; of cap, covered with white bloom, easily rubbed off
Gleba -- the spore mass of a Gasteromycete
Globose -- spherical, like a globe; of spores, spherical with round outline, according to one set of criteria ratio of length to width is 1.01-1.05: a greater ratio would indicate subglobose, elliptic etc.
Gloeocystidia (plural gloeocystidium) -- thin-walled cystidium with granular or oily contents; sometimes synonymous with chrysocystidium
Gluten -- the dissolved gelatinous hyphaeof certain tissues
Glutinous -- slimy, having a highly viscid gelatinous layer, more than viscid
Granular -- same as granulose
Granulose -- covered with granules
Greasy -- slippery or oily but not viscid (sticky) or slimy, same as lubricous
Gregarious -- growing in close groups but not tufted or clustered, A growth form in which mushrooms fruit in relatively close proximity.

Group -- a cluster of taxonomically related similar species typified by a particular species, as in Conocybe tenera group, sometimes used of a group of similar-looking species
Guttate -- having drop-like spots; drop-shaped
Guttulate -- of spores, containing an oil droplet
Habit -- the general external and characteristic appearance of mushrooms, and manner in which they are found growing
Habitat -- the natural place of growth
Hair brown -- grayish brown
Hairy -- covered by an arrangement of fibrils or mycelial strands resembling hairs
Hallucinogen -- capable of producing disturbances in (usually visual) perception
Hardwood -- any tree that is not a conifer
Hazel -- light to moderate yellowish brown; the color of the shell of the ripe hazelnut
Herbaceous -- said of those flowering plants that die annually at least down to the roots (i.e. non-woody flowering plants)
Herbarium -- a collection of dried plants or fungi arranged systematically
Heterodiametric -- of spore sizes, the average length divided by the average width has a value greater than 1.28: with subisodiametric spores this value is 1.16-1.27, and with isodiametric spores it is 1.0-1.15
Heterogeneous -- composed of unlike tissues; composed of more than one cell type
Heteromerous -- of trama in Russulaceae, having sphaerocyst nests among ilamentous hyphae
Heteromorphous -- composed of more than one structure
Hilar appendage -- nipple-like projection on spore which corresponds to the area that was attached to the basidium, sometimes used to refer to a projection on the other end of the spore, same as apiculus and hilar appendix
Hilar appendix -- same as hilar appendage
Hirsute -- covered with long stiff hairs
Hispid -- covered with long rough hairs or bristles, coarser or stiffer than hirsute
Hoary -- covered with dense silky down; canescent; with a silvery sheen as if covered with frost
Hollow -- of stem, having the flesh empty of fibrils, same as fistulose or tubular
Holotype -- the single element on which the describing author based a name, in the case of a mushroom species, the collection on which the describing author based the name
Homogeneous -- the same throughout
Hooked -- of gill attachment, same as uncinate
Homogeneous -- composed of like tissues; composed of one cell type
Horizontal -- of gills, attached in straight line perpendicular to stem
Host -- plant or animal on or in which a parasitic fungus exists
Humicolous -- living in humus
Humus -- decaying organic material in or on soil
Hyaline -- colorless
Hygrophanous -- cap surface changing color markedly as it dries, usually having a water-soaked appearance when wet and turning a lighter opaque color on drying. Having the characteristic of changing color upon drying.
Hygrophoroid -- applied to any mushroom with thick wax-like gills, like Hygrophorus
Hymeniform -- resembling a hymenium in form
Hymenophore -- spore-bearing surface
Hypha (plural hyphae) -- threadlike fungal cell
Hyphidium -- a little, or strongly, modified terminal hypha in the hymenium (spore-bearing surface), distinguished as follows by Donk: haplo- (simple - ) unmodified, unbranched or little branched; dendro- (dendrophysis) irregularly or strongly branched; dicho- (dichophysis) repeatedly dichotomously branched; acantho- (acanthophysis) having pin-like outgrowths near apex; synonymous or near synonymous are paraphysis, pseudoparaphysis, paraphysoid, dikaryoparaphysis and pseudophysis sensu Singer 1962. A little, or strongly, modified terminal hypha in the hymenium (spore-bearing surface)
Hypochnoid -- having effused, resupinate, dry, rather loosely intertwined hyphae
Hypoderm -- a differentiated region just below the pileipellis or stipitipellis, in most instances the same as the subpellis
Hypogeous -- fruiting underground
Hyphoid -- like hyphae; cobwebby
Hymenium -- The spore-bearing surface of a mushroom is called the "hymenium". The tubes make up the hymenium of a mushroom with pores. The gills constitute the hymenium of gilled mushrooms; In morchella(morels), false morels, helvella and similar mushrooms, the hymenium is the cap surface. The hymenium is covered with microscopic basidia or asci, which hold spores until they are released. Fertile area of fruiting body where spores are produced (in gilled mushrooms the surface of the gills), or the surface cell layer that produces the spores
Imbricate -- each growing just above the others, as with roof shingles
Inamyloid -- remaining clear or becoming yellow in Melzer's reagent, not amyloid or dextrinoid, same as nonamyloid
Incarnate -- flesh-colored
Incrassated -- widened
Incrusted -- covered with a thin, hard crust; of hyphae, with matter located on their outer wall; of cystidia, covered with crystalline or amorphous deposit, particularly at the top
Incurved -- of cap margin, curved inwards toward stem, but less than inrolled
Inequilateral -- of spores, means that a line drawn through the length of spore does not divide equal mirror images
Inferior -- of ring, located near base of stem, or at least below the middle
Inflated -- enlarged in some part; of a cell, enlarged at either the tip, middle or base
Inflexed -- bent inward, incurved
Inflorescence -- the arrangement of flowers on a floral axis
Infundibuliform -- funnel-shaped
Ingrown -- of stem, same as insititious and ingrown
Ink cap -- Common name of the Coprinus genus-- the caps become an inky mess when aged.
Innate -- usually of fibrils of scales, meaning that they are not raised from the surface or readily removed from it
Inrolled -- of cap margin, rolled inwards so that the edge of the margin is actually points toward gills
Inserted -- of stem, same as insititious and ingrown
Insititious -- of stem, devoid or any fibrils or hyphae at point of attachment to substrate, same as inserted and ingrown
Intercalary -- interposed, interpolated, placed in between
Intermediate -- of a ring (annulus), part way between a hanging or (skirtlike) ring which flares downward, and a sheathlike ring which flares upward, also referred to as a level ring
Intermediates -- subgills
Interveined -- of gills, connected by "veins" (ridges) that run between gills
Intervenose -- of gills, connected by "veins" (ridges) that run between gills
Interwoven -- hyphae entwined or tangled, not forming a regular pattern
Involute -- rolled in: in the case of the cap margin, rolled down and in, similar to inrolled
Isabelline -- color of unbleached linen, dingy yellowish brown, pale tan, similar to alutaceous, in Ridgway 1912 isabella color is a yellow brown
Isodiametric -- radially symmetrical; of spore sizes, according to one set of criteria the average length divided by the average width has a value less than 1.16: with subisodiametric spores this value is 1.16-1.27, and with heterodiametric spores it is greater than 1.27
Isotype -- a duplicate or part of the type collection, other than the holotype
Ixo -- a prefix indicating that elements are gelatinized
Ixocutis -- a curtis (with hyphae appressed to surface) embedded in slime
Ixolattice -- may be a well-developed ixocutis in which the hyphae have become very widely separated by slime, and the arrangement presented is modified to that of a tangled mass of hyphae, or may be an ixotrichodermium that collapses to form an ixolattice
Ixotrichodermium -- a trichodermium with slime present in it
Kaiser brown -- deep ferruginous
KOH -- potassium hydroxide, an agent commonly used to revive dried mushroom material, or show chemical reactions on the surface of the mushroom, or chemical reactions under the microscope. The chemical Potassium Hydroxide. Used in a 3% solution, it is a standard mounting medium used to rehydrate material for microscopic examination. It may also be used as a macro- or micro-chemical reagent differentially staining the tissues of some species. Concentrated solutions of this chemical are caustic and should be handled with care.
Kingdom -- one of five groups of living organisms: Monera (including bacteria and blue-green algae), Protoctista (including protozoans, most algae and three phyla of fungi), Animalia (animals), Plantae (plants), and Eumycota (the rest of the fungi)
Laccate -- looking as though varnished
Lacerate -- irregularly torn
Laciniate -- of margin or cap or annulus, cut more coarsely than fringed, slashed
Lactiferous -- of hyphae in flesh, bearing a milky juice
Lacuna (plural lacunae) -- hole or hollow
Lacunose -- having holes or hollows
Lageniform -- of cystidia, swollen at the base with the middle and top part tapered into a long beak, like a gourd, therefore gourd-like
Lamella (plural lamellae) -- gill, the spore-bearing platelike structure extending underneath and from the center of the cap
Lamellae -- The technical term used to describe the gills of a mushroom.
Lamellula (plural lamellulae) -- the short gills that do not span the entire distance from margin to stem
Lamprocystidium (plural lamprocystidia) -- thick-walled cystidium
Lanate -- same as woolly
Lanceolate -- like a lance, many times longer than broad, and tapering; of cystidia somewhat wider in middle and tapered at both ends
Lateral -- of a stem, attached to the side of the cap
Latericeous -- brick-red
Latex -- juice or milk of a Lactarius; juice usually of a milky color but also applied to other colors
Lectotype -- an element selected in a later work from the original material where no holotype was designated
Lens -- a hand magnifying glass
Lenticel -- any of the raised pores in the stems or branches of woody plants that allow gas exchange between the atmosphere and the internal tissues
Lentiform -- shaped like a lens
Leptocystidia -- smooth thin-walled cystidia
Lecythiform -- of cystidia, wide at base with middle tapered into narrow neck and top swollen into a head, like a bowling pin (lecythiform refers to a Greek stoppered bottle)
Lichen -- a dual organism in which a fungus (usually an ascomyce ascomycete but occasionally a basidiomycete) maintains a green alga or cyanobacterium captive for mutual benefit
Lignicolous -- living in, on, or out of wood. Living on wood.
Lignin -- the organic substance of woody tissue other than cellulose
Lilac -- the color of flowers of the lilac shrub, a pale purple or mauve
Liver -- brown deep reddish brown
Livid -- a dark blue-gray color
Lobed -- with rather large, rounded divisions on the margin
Long decurrent -- refers to decurrent gills that proceed a long way down the stem, for instance proceeding further down the stem than the height of the gills
Lubricous -- greasy or slippery or oily but not viscid (sticky) or slimy
Luteous -- dull egg-yellow
Lutescent -- becoming luteous
Macrofungi -- Fungi visible to the naked eye.
Margin -- The outermost edge of the gill ( as in the edge facing downwards)
Macrocystidium (plural macrocystidia) -- cystidium arising deep in the flesh of Lactarius or Russula; any large cystidium
Macromorphological -- concerning structure that can be seen with the naked eye
Macroscopic -- visible to the naked eye, without a microscope
Maculate -- spotted
Micron -- A metric unit of measure equal to one one-thousandth (1/1000) of a Millimeter. Written as 0.001mm.
Milk -- A liquid exuded from certain species such as Lactarius. See Photo Below.



Mycelium -- The vegetative part of the fungus which grows in the host or soil and produces the fruit body. The mycelium is like a mass of often microscopic fibers. It is underground and may be invisible. The visible mushroom is like it's fruit, the mycelium being the fruit tree.
Mycology -- The Study Of Fungi.
Mycorrhiza -- A symbiotic relationship between a fungus and the roots of a plant.Nomen provisorium -- a name proposed provisionally, not yet an official name
Nonamyloid -- remaining clear or becoming yellow in Melzer's reagent, not amyloid or dextrinoid, same as inamyloid - distinguish from "not amyloid" which would include nonamyloid and dextrinoid
Notched -- refers to gills that are uncinate or sinuate or emarginate, as if a wedge of gill had been removed near the stem: if the line of the bottom edge of the gill curves down sharply, gills are uncinate, if it curves gradually toward the stem reaching it more or less horizontally, gills are sinuate (emarginate)
Nutant -- nodding
Obclavate -- club-shaped in the opposite direction to that expected; of cystidia, with base swollen and narrowing at middle and top
Obconic -- like an ice-cream cone with point down
Obligate -- invariably found in a particular situation, usually in reference to organisms that must live in a particular association with another
Oblong -- of spores, elongated with approximately parallel sides; according to one set of criteria, ratio of length to width is 1.6-2: shorter would be elliptic and longer cylindric; however, spores in this range are often referred to as narrowly elliptic
Obovate -- ovate with the larger end in the opposite direction to the usual
Obovoid -- ovoid with the larger end in the opposite direction to the usual
Obpyriform -- pear-shaped in the opposite direction to the usual one
Obsolete -- (of annulas, scales etc.) very imperfectly developed, hardly perceptible; of terms, no longer in use
Obtuse -- blunt, not pointed; greater than a right angle
Obtusely conic -- rounded or blunt cone-shaped
Obtusely umbonate -- broadly umbonate, not with sharp umbo
Ochraceous -- ochre-yellowish, yellow-orange with a brownish tinge
Ochraceous-buff -- a very pale but dingy yellow
Ochraceous-tawny -- like the color of a dingy or dirty lion
Ochre -- between warm buff and yellow to brownish orange
Ochreate -- of volva, sheathing the stem at base like a stocking
Odor -- The smell of the fruiting body
Olivaceous -- olive gray-brown; with an olive shadeO
Omphalinoid -- of general form of the genus Omphalina, with broadly convex to depressed cap, decurrent or subdecurrent gills, cartilaginous stem, and no ring or volva
Opaque -- not transparent or translucent, often used of cap margin where gills do not show through as striations
Orbicular -- circular
Order -- a classification grouping below class but above family, genus and species: suffix is -ales
Organism -- individual living bacterium, protozoan, animal, plant, fungus etc.
Ornamentation -- any projections outside the structural surface such as fibrils, tomentum, hairs, warts, scales, spines, ridges, etc.
Ornamentation type -- Types of Russula spore ornamentation have been designated by Singer (1932), using Roman numerals, Pearson, (P1-P11), Rayner (15 types), and Dave Patterson in his Key to the Eastern U.S. Russulas (A1 to E3). The last are used in this program. For an illustration of the Patterson types, use the Glossary on the List menu.
Outer veil -- same as universal veil
Oval -- like the outline of an egg
Ovate -- similar to oval but some regard as more pointed at the narrower end
Ovoid -- shaped like an egg, same as oval, but sometimes implying 3-dimensional shape
Palisadoderm -- type of pellis in which terminal elements reach the same level and form a palisade of inflated somewhat elongate cells
Pallid -- very pale in color, almost a dull whitish
Papilla (plural papillae) -- a small nipple-like protuberance
Papillate -- with papilla or papillae on surface
Parabolic -- of cap, with the height greater than the width, the top rounded
Parallel -- of hyphae, arranged more or less parallel to each other
Parasitic -- feeding on another living organisms; living at the expense of other organisms to their detriment
Parasitize -- feed on another living organism
Pallid -- Pale, light in color.Parasitic -- A fungus that grows at the expense of another organism, drawing nourishment from it. Example: Armillaria mellea (also called the honey mushroom or oak root rot fungus).
Partial veil -- The covering of the gills while very young that breaks open, often leaving remnants on the stem (stipe). Inner veil of tissue which joins the stem to the cap edge at first in some species of mushrooms, and often breaks to leave a ring on stem and remnants hanging from the cap margin; partial veils are usually either membraneous or cortinateSee Photo Below.


Partial Veil Remnants -- The partial veil may also leave fragments clinging to the margin of the cap. Sometimes, these partial veil remnants are the only evidence one has that there was ever a partial veil at all, and their presence or absence is frequently important in mushroom identification. See Photo Below.


Patches -- Fragments of universal veil tissue left on the cap of a mushroom after it has matured are called "patches." Like warts, patches are easily washed off by rain. Patches are similar to warts, but there are fewer of them, and they normally involve larger fragments. Please see the below photo.

Parenchymatous -- fundamental tissue of plants composed of thin-walled cells able to divide; organ tissue as opposed to connective tissue
Patterson ornamentation types -- Types of Russula spore ornamentation as designated by Dave Patterson in his Key to the Eastern U.S. Russulas. For an illustration of these types, use the Glossary on the List menu. Other classifications of Russula spores have been made by Singer (1932), using Roman numerals, Pearson, (P1-P11), and Rayner (15 types)
PDAB -- a solution of p-diaminobenzaldehyde in 70% ethanol
Pecan brown -- orangy pinkish brown
Pectinate -- (of cap margin) resembling the teeth of a comb; same as striate
Pedicel -- of cystidia, a slender stalk
Pellicle -- an upper surface layer on cap surface that can undergo gelatinization, making the cap viscid (sticky) to the touch; often it can be peeled away from the cap, may be thought of as covering cuticle; same as cuticle or as thinner and more definite
Pellis -- surface layer of cells, same as "cuticle", if the pellis is one layer it is called a suprapellis, if 2 layers, the outer is the suprapellis and the inner the subpellis, if 3 layers, the middle is called the mediopellis
Pellucid -- translucent
Pendant -- hanging down, skirt like
Peppery -- with acrid taste, giving the tongue a burning sensation
Peridium -- the outer covering enclosing the spore mass in gastermycetes
Peronate -- sheathlike; booted
Petaloid -- shaped like the petal of a flower (narrowed somewhat at base), similar to spathulate
Phaseoliform -- bean-shaped
Phylogeny -- the history of the evolution of the group to which a species belongs
Phylum (plural phyla) -- classification grouping below kingdom but above class
Phenolic -- Having an odor of the chemical phenol.Pileus -- The cap of a mushroom. The hymenium-supporting part of agarics.
Pileate -- having a cap
Pileipellis -- the outer cellular layer of the cap (pileus), excluding veils, used in microscopic descriptions: it may be undifferentiated from the underlying tissue, or arranged parallel to surface (curtis) or arranged perpendicularly to surface (derm)
Pileocystidium (plural pileocystidia) -- sterile cell (cystidium) on the surface of the cap
Pilocystidium -- same as pileocystidium
Pilose -- (of cap), covered with long, soft, hairy filaments
Pip-shaped -- (of spores), shaped like an apple seed; sometimes used to describe spores with a plage at one end that would be described as elliptic by other authors
Pith -- the central stuffing of some mushrooms
Pitted -- with small depressions
Plage -- a distinctive flattened area on the dorsal side of the spore (the side facing the central axis of the basidium on which the spore develops), near the hilar appendage (the part of the spore that was attached to the basidium), also known as suprahilar disc; if plage amyloid it is known as a hilar spot, if depressed as suprahilar depression
Plane -- having a flat surface; of cap, having a horizontal more or less flat surface, with the margin on the same level as the center, same as flat or applanate
Plano-convex -- a convex cap with a flattened disc, same as broadly convex
Plectenchyma -- a thick tissue formed by hyphae becoming twisted and fixed together; it is prosenchyma (proso-) when the hyphal elements are seen to be hyphae and pseudoparenchyma (para-) when they are not
Pleurocystidium (plural pleurocystidia) -- a sterile cell (cystidium) located on the face (side) of a gill
Pleuropseudocystidium (plural pleuropseudocystidia) -- pseudocystidium located on the face (side) of a gill
Pleurotoid -- resembling in general form the genus Pleurotus, may be applied to any gilled mushroom either without a stem or with a stem attached in a lateral or off-center manner
Pliable -- capable of bending, easily flexible
Pliant -- being pliable without breaking, flexible, not rigid or firm
Plicate -- folded like a fan, pleated
Plumbeous -- lead-gray
Pluteotoid -- resembling in general form the genus Pluteus, with free or finely adnexed gills, lacking a ring or volva
Pocket rot -- a rot producing hollow pockets in a tree
Polar view -- end view of a spore
Polycystoderm -- same as epithelium
Polymorphic -- with many forms or shapes
Polypores -- the shelf or bracket fungi which produce spores on the inside of vertically oriented tubes (ending in pores) that do not separate easily from cap and are often tough, generally in Order Poriales. Common name of tough-textured fungi with pores
Pores -- Pores are very small holes on the underneath side of some mushroom species such as Boletes as opposed to gills. They produce their spores on the inside surfaces of tubes. A circular depression in place of the gill of many non-gilled species; a circular depression on the spores of many species: see germ pore


Powdery -- looking finely powdered or very finely granular, sometimes used here as equivalent to pruinose
Primordium -- the earliest stage of development
Profile view -- of spores, when the basidium with attached spores is viewed from the side, the spores directly above and in the center are being viewed in face view and the ones at the sides are in profile view, when not attached to basidium the shape that is wider will generally be the face view if there is a difference
Protean -- extremely variable
Prout's brown -- dark yellow-brown, warm mid-brown
Pruinate -- same as pruinose
Pruinose -- looking finely powdered or finely granular, often due to cystidia projecting from surface
Pseudoamyloid -- same as dextrinoid
Pseudocystidium (plural pseudocystidia) -- cystidiumderived from a conducting element, filamenteous to fusoid, oily contents, embedded or not projecting
Pseudoparaphysis (plural pseudoparaphyses) -- elements found in certain gilled mushrooms that are similar to the paraphyses found as sterile elements among the asci on spore bearing surfaces of ascomycetes
Pseudoparenchyma -- thick tissue formed by hyphae becoming twisted and fixed together, in which the hyphal elements are not seen to be hyphae
Pseudorhiza -- a long rootlike extension of the lower stem
Psilocin -- 4-hydroxy-N,N-dimethyltryptamine, a hallucinogenic substance found in some species of Psilocybe, Panaeolus, Conocybe, Gymnopilus, Inocybe, and Pluteus, giving a bluing reaction in the tissue of a mushroom as it breaks down
Psilocybin -- O-phosphoryl-4-hydroxy-N,N-dimethyltryptamine, a hallucinogenic substance found in some species of Psilocybe, Panaeolus, Conocybe, Gymnopilus, Inocybe, and Pluteus
Puberulent -- minutely pubescent = puberulous = pubescent
Puberulous -- minutely pubescent = puberulent = pubescent
Pubescence -- a covering of soft short downy hairs
Pubescent -- covered with soft short downy hairs, but "downy" may differ by having slightly larger hairsPulverulent -- powdery
Pulvinate -- cushion shaped
Punctate -- marked with dots consisting of hollows, depressions, spots, raised-joined scales, or agglutinated fibrils, all very small. Having minute scales or points on the surface.
Putrescent -- in the process of rotting or smelling as if it is. Tending to decay rapidly.
Pyriform -- pear-shaped
Quaker drab -- medium gray
Radicating -- forming a root
Radicate -- with a root or pseudorhiza
Rameales structure -- type of pellis in which the hyphae of the outer layer are basically prostrate but possess short vertical branches, or the hyphae branch irregularly as in some species of Marasmius, Coprinus, Marasmiellus, or Xeromphalina
Raphanoid -- like radish
Raw sienna -- brownish yellow-orange, bright yellow brown
Receptacle -- any hymenium-supporting structure
Recumbent -- hangs down, rests on surface from which it extends
Recurved -- curved back: when used of cap margin or scales means curved back upward
Reflexed -- turned back to form a cap; turned up or back
Regular -- of gill tissue, same as parallel
Reniform -- kidney-shaped
Repand -- (of cap) wavy on margin and turned back or elevated
Repent -- of hyphae, prostrate, lying flat
Refractile -- having power to refract (i.e. to change the direction of light)
Remote -- (of gills) proximal end free and at some distance from the stemResupinate -- lying flat on what it is growing on, without a stem or well-formed cap, with the gills or pores facing outward
Reticulate -- Usually referring to a fish-net or crosshatch pattern. See Photo Below. Covered with a network of interlacing lines

Reviving -- said of fruiting body which shrivels in dry weather or when dried and takes on its natural shape when wet
Revolute -- (of cap margin), rolled back or up
Rhizoid -- root-like structure from the base of fruiting body
Rhizome -- a rootlike subterranean stem commonly horizontal in position that usually produces roots below and sends up shoots progressively from the upper surface
Rhizomorph -- cordlike strand of twisted hyphae present around base of stem, often dark colored
Rhizomorphs -- Mycelial cords or strands, called "rhizomorphs," are found in some species, and their presence often helps in identification decisions.
Ring -- Remnant of the veil on the stem just under the cap, see photos. Annulas, collar of tissue on stem formed by ruptured of the veil that initially joins the stem to the cap edgeSee annulus.


Ring Zone -- A "ring zone" is a zone on the upper part of the stem resulting from the deterioration of the partial veil. Ring zones are frequently not as prominent as the rust-colored zone on the Cortinarius species shown in the photo below, they may be barely noticeable.

Rhomboid -- having or nearly having the shape of a rhombus; a parallelogram with angles that are not right angles, and unequal adjacent sides
Rhombus -- an outline with four equal sides, but with angles that are not right angles
Ridged -- of spores, with narrow raised straight or curved strips on the surface of the spore
Rimose -- cracked, referring to surface of cap or stem
Rimulose -- finely cracked
Rivulose -- arranged like rivulets in the soil, marked with riverlike lines
Rood's brown -- dark pinkish brown
Rostrate -- with a beak; of cystidia, having a beak-like or finger-like protuberance called a rostrum
Saccardo's umber -- close to date brown, dark yellow brown
Saccate -- of a volva, shaped like a sac, cup or sheath
Salmon-buff -- a dingy pink
Sanguineous -- blood-red
Saprobe, Sabprobic, Saprophytes -- Fungi that receive nourishment from dead organic material. -- These are Mushrooms that are saprobes survive by decomposing dead or decaying organic material. Many wood rotting fungi are saprobes, and help decompose dead wood--but other wood rotters are parasitic and attack living wood. Saphrophytic organism
Saprophytic -- living on decaying organic matter
Sarcodimitic -- consisting of generative hyphae and thick-walled, long, inflating fusiform elements, as in Gerronema
Sayal brown -- close to cinnamon in color, dull to dark cinnamon; between a moderate orange and a moderate yellowish-brown
Scaber -- short projecting scale or tufted hairs
Scabrous -- Having conspicuous scales on the surface. Roughened by short projecting rigid scalesSee Photo Below.

Scale -- piece of tissue on surface that is not especially elongated, differentiated from surface by color or by projecting from it
Scarlet -- a brilliant red color slightly tinged with orange
Sclerotium -- a knot or firm frequently rounded mass of hyphae, usually underground, sometimes giving rise to mycelium or a fruiting body
Scrobicula (plural scrobiculae) -- a large conspicuous shallowly sunken spot, pit, hollow, or depression
Scrobiculate -- having scrobiculae, pitted with conspicuous wet-appearing slightly depressed areas
Scum-like -- of cap, having an area of superficial dark-colored fibrils
Scurfy -- surface covered with branlike particles resembling scales, same as furfuraceous
Seceding -- refers to gills that have separated in their attachment to the stem and have the appearance of being free, often leaving longitudinal lines on the stem where the gills were once connected
Secondary spores -- not borne on basidia, conidia, chlamydospores etc., formed directly on the mycelium or on hyphae of the fruit body
Secotioid -- suggesting an undeveloped or aborted gilled mushroom, resembling Secotium
Senescent -- becoming old
Sensu -- in the sense of
Sensu lato -- in a wide sense
Sensu stricto -- in a narrow sense
Separable -- said of stem or gill easily removed from cap
Separate -- of gill attachment, same as secede
Sepia -- a moderate brown; a brownish gray to a dark olive-brown
Septate -- partitioned with cross-walls
Septum (plural septa) -- cross-wall in hyphae
Sequestrate -- describes fruiting bodies that have evolved from those that forcibly discharge spores to a closed or even underground form in which spores are retained until it decays or is eaten by an animal, the word referring to spores which have been sequestered (hidden). Lactarius is thought to give rise to Arcangeliella (mostly above ground, but gills not exposed or vertically oriented and do not discharge spores forcibly) and Zelleromyces (underground, no true stem). Russula is thought to give rise to Macowanites (mostly above ground), Gymnomyces (underground, no stem), Elasmomyces (no sphaerocysts in hymenial tissue), and Martiella (no sphaerocysts in hymenial tissue, underground without stem). Cortinarius is thought to give rise to Thaxterogaster (above ground) and Hymenogaster (underground, no stem). Agaricus is thought to give rise to Endoptychum and Longula. Chroogomphus is thought to give rise to Brauniellula (often buried or half buried). Pholiota is thought to be related to Nivatogastrium (grows on wood). Other postulated sequestrates are given in brackets: Amanita (Torrendia), Bolbitiaceae (Gastrocybe), Boletus (Gastroboletus), Coprinus (Podaxis), Entoloma (Richonia), Gomphidius (Gomphogaster), Lepiotaceae (Notholepiota), Paxillaceae (Austrogaster, Gymnopaxillus), Strobilomycetaceae (Gautieria), Suillus (Rhizopogon, Alpova, Truncocolumella, Gastrosuillus).
Sericeous -- silky, like silk
Serrate -- saw-toothed to almost ragged
Serrulate -- finely serrate
Sessile -- lacking a stem
Seta (plural setae) -- pointed, elongated, thick-walled sterile cells
Setula (plural setulae) -- a fine bristle; a thick walled, pigmented, terminal element of a tramal cystidium
Shaggy -- rough as with long hair or wool
Sheathlike -- of a ring, clinging to the stem and opening upwards
Short decurrent -- refers to decurrent gills that do not proceed down stem far: for instance, as much as the width of the gills
Siderophilous -- of basidia, with granules that darken when heated in acetocarmine
Sienna -- raw sienna is brownish yellow-orange or bright yellow brown; burnt sienna is a watery strong red-brown or dark orange brown
Sinuate -- of gill attachment, refers to gills with a lower edge that curves up close to the stem then curves back to reach the stem more or less horizontally; of cap margin means wavy or undulating. Referring to a type of gill attachment, specifically gills that are notched at their point of attachment to the stipe. See Photo Below.

Sinuous -- crooked or curved
Skeletal hypha -- thick-walled, little branched non-septate hypha
Skirtlike -- of a ring (annulas), hanging down like a skirt
Slimy -- having a thick layer of slime, more than viscid or glutinous
Smooth -- of a surface, without projections, often equivalent to bald or glabrous; but may be described as bumpy and bald, or finely powdery and smooth; of cap margin may mean not wavy or lobed, or may mean not grooved; of spores, not spiny rough, or ridged
Snuff-brown -- same as tobacco-brown, a dark sepia, a dull yellowish brown, a dull cinnamon brown
Solid -- not hollow; feeling hard
Solitary -- not growing in the immediate neighborhood of other individuals
Sordid -- dingy-looking
Sp. nov. -- new species
Sphagnum -- a genus of moss that grows in bogs
Sphaerocyst -- Round swollen cells usually formed in clusters, characteristically found in the Russulaceae. Sphaerocysts make the flesh of Russulas brittle.
Spathulate -- shaped like a spatula or spoon, oblong with a narrowing base
Species -- classification grouping below family and genus, often used for organisms capable of interbreeding (though less common "hybrids" can occur between species), among anamorphic fungi that are not known to breed sexually, it refers to a certain level of similarity in form or function; named by genus name in upper case and species name in lower case, e.g. Russula emetica
Spermatic -- resembling the odor of human sperm or semen
Sphaerocyst, spherocyst -- a round or swollen cell in flesh of certain mushrooms, particularly Russula and Lactarius
Sphaerocyte, spherocyte -- round cell of pellis or veil
Species -- A group of individuals with certain common characteristics
Spore Print -- The spore material left on paper or under a mushroom in the wild when the cap is left for a period of time -- the color can be an important identifying characteristic. See Photo Below of a natural occurrence. Notice the spore deposit on the leaf. A visible deposit of spores obtained by allowing a gilled mushroom to drop spores onto white paper for a few hours or overnight

Stem -- See stipe.
Stipe -- The technical name for a mushroom stem or stalk. The stipe supports the pileus (cap) in the agarics (gilled mushrooms).

Spine -- long slender sharp projection
Spiny -- having many spines
Sporadic -- irregular in its occurrence, either in time or location
Spore -- reproductive cell or "seed" of a fungus, produced on specialized cells, which in gilled mushrooms are on the gills. Reproductive cell found in fungi. A microscopic part of the fungus which can germinate to reproduce the fungus
Spore wall -- in the most complex spore wall there are five layers from outer to inner: perisporium, non-pigmented and usually enveloping spore like a bag which may disappear; exosporium, usually non-pigmented and can often be distinguished chemically from other layers, episporium, a continuation of outer wall of basidium, the thickest layer and the one providing structural support, mesosporium, a barely distinguishable delicate structure, and endosporium, which can vary from very thick (in which case it can then be divided into inner and outer part) or seemingly absent, or truly absent; the presence or absence of layers varies with species
Sporocarp -- a structure in which or on which spores are produced, often used for fruiting body, consisting of cap, gills, and stem
Spotted -- with roundish areas different in color from the rest of the surface
Squamose -- scaly, with moderate to large scales
Squamule -- scale
Squamulose -- with small scales
Squarrose -- covered with upright or curved-up pointed scales
Squarrulose -- covered with small upright pointed scales
Stalk -- same as stipe or stem
Stature -- characteristic shape
Stellate -- star-shaped
Stem -- the column supporting the cap in most mushroom, more correctly called the stipe
Sterigmata -- elongated appendages or "arms" on the basidium upon which spores are borne
Sterile -- not producing spores
Stipe -- the correct name for the "stem" of a mushroom
Stipitate -- having a stipe (or stem)
Stipitipellis -- surface layer of the stem
Stramineous -- straw-colored
Strangulate -- constricted
Streaked -- having faint lines or bands, used when appressed fibrils appear like bands or faint lines
Stria (plural striae) -- lines or fine grooves which may be parallel or radiating
Striate -- marked with lines or fine grooves which may be parallel or radiating
Strigose -- having long stiff hairs
Stroma (plural stromata) -- a mass or matrix of vegetative hyphae, with or without tissue of the host, sometimes sclerotium-like in form, in or on which spores or fruiting bodys are produced
Stuffed -- containing loose material in the interior, not hollow or solid
Sub- -- near, nearly, more or less, somewhat, slightly; below or under; subdivision of
Subclose -- a term used occasionally of gill spacing, intermediate between close and crowded, might also be used to mean more or less close
Subcrowded -- a term used occasionally of gill spacing, intermediate between close and crowded, might also be used to mean more or less crowded
Subdecurrent -- of gills, meaning short decurrent or nearly decurrent or somewhat decurrent (i.e. intermediate between adnate and decurrent, when attachment extends slightly further down stem than when adnate)
Subdistant -- of gill spacing, intermediate between close and distant, the order being crowded, (subcrowded), (subclose), close, subdistant, distant
Subfusiform -- of spores, elongated, tapered at one end and rounded at the other
Subgills -- the short gills that do not span the entire distance from margin to stem
Subglobose -- of spores, nearly spherical or round; according to one set of criteria (Bas' work on Amanita) ratio of length to width 1.05-1.15
Subhymenium -- a differentiated tissue just beneath the hymenium
Subicule -- same as subiculum
Subiculum -- a net-, wool-, or crust-like growth of mycelium under fruiting bodies
Subisodiametric -- of spore sizes, the average length divided by the average width has a value from 1.16-1.27: with isodiametric spores this value is 1.0-1.15, and with heterodiametric spores it is greater than 1.27
Subpellis -- the layer that separates the pellis from the trama; often considered the same as the hpoderm
Substrate -- the material that a fungus is growing on
Substratum -- substrate, the material that a fungus is growing on
Subtomentose -- with a somewhat dense layer of matted down or soft hairs; or like a newly sheared lamb
Subulate -- awl-shaped (subula = awl); of cystidia, swollen between the middle and the slightly tapered base and pointed at top
Subviscid -- slightly sticky, thinly viscid
Sulcate -- grooved, furrowed
Superior -- of a ring, forming on the upper part of the stem
Suprahilar disc -- same as plage
Suprapellis -- the outermost layer of the pellis, it may be undifferentiated (cells not distinct from those of underlying flesh), derm (cells arranged perpendicularly to surface), or cutis (cells arranged more or less parallel to surface)
Synonym -- another name for the same species, especially an earlier or illegitimate name not currently used for the species; if two or more names are based on the same type, they are homotypic synonyms, sometimes indicated by three horizontal lines between the two names, but if they are based on different types, they are heterotypic, sometimes indicated by two horizontal lines between the two names; in this program, alternate names following the primary name are earlier or later or illegitimate names representing all or part of the concept of the primary name: the primary name includes the alternate name, but the alternate name may not include the whole taxon represented by the primary name
Synonymous -- representing the same species
Synonymize -- subsume a species name under another species name
Tacky -- slightly sticky but not truly subviscid or viscid
Tan -- leather-colored, similar to undressed leather
Taproot -- a root which grows vertically downward, narrowing from top to bottom
Tawny -- approximately the color of a lion, between yellow brown and rusty brown; used by some as more orange, fox-colored, equivalent to fulvous
Taxon (plural taxa) -- a named form, variety, species etc.
Taxonomy -- The classification of organisms to show relationships to other organisms.
Tenacious -- tough
Terrestrial -- appearing to grow from the ground, or on the ground, as opposed to growing on wood
Terete -- of stem, cylindric or rounded; not compressed or irregular
Testaceou -- brown mixed with yellow or red, close to brick color, in Ridgway 1912, an orangy pink
Thick -- term used for width of stem, depth of cap flesh, or the distance between the faces of one gill
Tibiiform -- of cystidia, somewhat ventricose (wider in middle) with long narrow neck and apex swollen into a head, supposedly like the tibia bone
Tier -- in reference to subgills, group of subgills, interspersed with gills usually at regular intervals, each tier being of roughly a certain length
Tilleul buff -- pallid or whitish
Tissue -- a group of hyphae which are similar in shape or form
Toadstool -- a mushroom, especially a poisonous one
Tobacco brown -- the color of tobacco as it is found in a cigar or cigarette
Tomentose -- covered with soft hairs, often soft densely matted hairs, like a woollen blanket
Tomentous -- same as tomentose
Tomentulose -- covered with short fine hairs or fibrils, which may be matted like a thin woollen blanket or erect according to different authors' interpretations; nearly tomentose but less than subtomentose
Tomentum -- a covering of densely matted woolly hairs
Tomentose -- Having very minute fine hairs on the surface. See Photo Below.

Toothed -- serrate on the edges; toothlike on the edges; of gills, with toothlike edges or decurrent by a short tooth
Tough -- strong, able to resist stress
Trama -- the tissue under the surface cell layers of cap, stem, or gills, usually referring to the flesh (context) as seen through the compound microscope
Translucent -- transmitting light diffusely, semitransparent
Translucent-striate -- refers to a cap that allows some light to pass through and which, as a result, shows the gills as darker radiating lines in the translucent area
Trichoderm -- a "cuticle" with hair-like elements projecting from the surface under the compound microscope, more or less perpendicularly, forming a turf, the individual elements unequal in length
Trichodermium -- same as trichoderm
Tricholomatoid -- resembling in general form the genus Tricholoma, with notched gills, fleshy-fibrous stem, and no ring or volva
Trimitic -- consisting of three kinds of hyphae: generative, binding and skeletal
Troops -- hundreds to even thousands of fruiting bodys growing within a few square yards
Trullisate -- resembling a small planting scoop
Truncate -- larger portion ending as if cut off, having the end square. Having a flattened or chopped off end like the end of a baseball bat.
Tubes -- See Pores.
Tuber -- a fleshy, lump-like or root-like stem base
Tubercle -- any wart-like or knob-like protuberance
Tuberculate -- with low bumps
Tuberculate-striate (or tuberculous-striate) -- of cap margin, furrowed radially with small bumps on the ridges
Tubular -- of stem, having the flesh empty of fibrils, same as fistulose or hollow; of hymenophore, composed of tubes, the opening of which is called a pore
Tufted -- as used here, the same as caespitose; may also be used to mean a small cluster or stems clustered with a common base, see clustered, caespitose, connate
Tuning fork basidia -- basidia of the jelly fungus order Dacrymycetales, shaped like a Y or a tuning fork
Turbinate -- top-shaped; of cystidia, swollen at top, tapered from middle downward, becoming abrupt at base
Type -- the element on which the descriptive matter fulfilling the conditions of valid publication of a scientific name is based; in the case of mushroom species, the collection of fruiting bodys from which the original concept of the taxonomic group (e.g. family, genus, species, variety, etc.) is derived
Type collection -- a collection of fruiting bodys from which the original concept of the taxonomic group (e.g. family, genus, species, variety etc) is derived
Umber -- a deep dull dark brown, smoky brown; earth brown sometimes with a very slight reddish tinge
Umbo -- a raised knob or mound at the center of the cap
Umbrinous -- olive-brown; umber
Umbilicate -- Having a small depression, e.g. as in a belly button. Refers to a cap with a narrow, moderate to deep depression in center which may or may not have a small umdo in the bottom


Universal veil -- Material which completely covers the young immature mushroom

Uncinate -- refers to gills with a lower edge that curves up as it comes close to the stem, then abruptly curved down to leave a "tooth" on stem, not proceeding further down stem than the imaginary line running straight along the lower gill edge to the stem, but sometimes used as equivalent to decurrent with tooth"
Undulate -- wavy
Unicolorous -- of one color
Universal veil -- the enveloping veil initially covering the whole mushroom including the top of the cap: when it breaks, it may leave fragments on the cap or the stem, or a volva at the base of the stem
Uplifted -- the margin of the cap turning upward
Upturned -- the margin of the cap turning upward
Urceolate -- having the shape of a pitcher, with a large body and small mouth
Urticoid -- with a swollen base and a long gradually narrowed apex
Utriform -- of cystidia, with a slight constriction below a large round head, like a bladder, therefore bladder-shaped
Vaginatoid -- applied to any mushroom with free or finely adnexed gills, a volva, and lacking an annulus
Velutinous -- velvety
Variety -- (abbreviated var.) a consistent appearing variation of a species, with more variation than a form, sufficiently hereditary as to characterize homogeneous populations
Variety -- (abbreviated var.) a consistent appearing variation of a species, with more variation than a form, sufficiently hereditary as to characterize homogeneous populations
Veil -- referring either to the partial veil which joins the stem to the cap edge at first, and often breaks to leave a ring on stem and remnants hanging from the cap margin, or the universal veil which initially covers the whole fruiting body including the top of the cap, always breaking and sometimes leaving fragments on the cap or the stem, or a volva at the base of the stem
Velar -- of the veil
Velum -- same as veil
Velutinous -- like velvet
Ventricose -- wider in the middle
Vermilion -- a bright red color with a strong orange tinge
Verrucose -- with warts; or with outgrowths smaller than if warted but larger than if verruculose (as used here, warty includes verrucose and verruculose)
Verruculose -- with moderate outgrowths smaller than if verrucose
Versiform -- with various shapes
Vesiculate -- of cystidia, with entire cell swollen or appearing inflated like a large sac or bladder (vesicle), with only the base abruptly tapered
Vesiculose -- same as vesiculate
Villose -- covered with long soft, weak hairs that collapse readily
Villose-tomentose -- having a densely matted layer of long, soft hairs which collapse readily
Vinaceous -- the color of red wine or red wine stains; a paler or grayish red; dull pinkish brown to dull grayish purple
Vinaceous-drab -- purple-gray
Violaceous -- of some violet hue
Virgate -- markedly streaked or striate, usually with dark-colored groups of fibrils, giving the appearance of bearing many small twigs
Viscid -- Slimy, sticky, viscous. Sticky but not slimy or lubricous: the mushroom usually feels somewhat slimy or slippery when wet but when dry may need to be wetted slightly to feel sticky; sometimes used to include slimy
Vitelline -- egg-yellow
Volva -- the remains o f the universal veil found at the base of the stem, usually in the form of a sac, collar or concentric rings. A sac-like structure formed at the base of a stipe, such as that found in Amanita species.

Warm buff -- pale yellowish buff
Warm sepia -- dark dull vinaceous brown, dark dingy cinnamon
Wart -- bumpy outgrowth found on caps, stems, and spores, which on caps and stems is generally somewhat wider than high
Warts -- Common in the Lepiota and Amanita species, see the photos below, notice the warts on the caps.



Waxy -- appearing as if coated with wax
Well-spaced -- referring to gills, corresponds to distant
White rot -- a rot that removes both lignin and cellulose
Wood brown -- dark avellaneous
Y-shaped basidia -- basidia of the jelly fungus order Dacrymycetales, shaped like a Y or a tuning fork
Zonate -- with circular bands of differing colors or ornamentation. Having a zoned appearance, usually referring to a mushroom cap that has concentric color bands that give it a zoned appearance. Common in Lactarius.
Zoned -- same as zonate
Is there a term that is not on this page? If so, please advise me so that I can add it! Thanks - Chris M
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